Virtual learning environments constitute current information systems' category for electronically supported training and development in (higher) education(al) and vocational training settings. Frequently expected advantages of using virtual learning environments refer, for instance, to the efficiency, individuality, ubiquity, timeliness and learning task orientation. However, a crucial precondition of realising such advantages is an appropriate system design. Hence, the question "Which specific design characteristics actually characterise successful virtual learning environments?" is of specific interest for training and development practice. This paper therefore discusses virtual learning environments' design characteristics by conducting an in-depth literature review. Based on this, a comprehensive set of diverse design characteristics of virtual learning environments as well as particular information associated with them are elicited, presented and discussed. Beyond this, particular implications for research and practice are derived. This may contribute to a successful development, implementation and (continuous) improvement/evaluation of virtual learning environments. (Contains 8 tables.)